# Kramer Lab #### Welcome to the Kramer Lab! ![Alpina Canadensis](/sites/g/files/omnuum6406/files/styles/hwp_28_10__1920x685/public/2025-08/alpina_canadensis_devseries_0.png?itok=LPdLjtY6) Welcome to the Kramer Lab! Our lab is very broadly interested in the evolution of floral morphology. We use molecular, morphological, and phylogenetic approaches to study how flowers have changed over the course of evolutionary time. Research projects in the lab cover a diverse set of topics, including gene lineage evolution and the effects of gene duplication, the morphological diversification of floral parts such as petals and fruits, and the evolutionary and ecological significance of pollinator interactions. Within this context, one of our major focus areas is the development of Aquilegia (columbine) as a new system for studying evolutionary and ecological questions, but we also work on other diverse lineages, including Loasaceae and Tropaeolum. As a lab, our goal is to create a work environment that is respectful, inclusive and equitable, and in which every individual feels safe to be their authentic self. We recognize that our research is enhanced by a diversity of thought and experiences and thus we welcome and actively seek out lab members from all backgrounds. As a group and as individuals, we seek to continually educate ourselves on systemic racism and how to combat it in academia and in our everyday lives. To these ends, we are actively engaged in outreach and antiracism efforts within Harvard and across the broader botanical community. We hope that these efforts will help address the chronic underrepresentation of diverse scientists in botany, which has been colored by a history of colonialism and exploitation. ## Latest News [### Congratulations to Teo on their acceptance to UC Santa Cruz! ](/news/2026/04/congratulations-teo-their-acceptance-uc-santa-cruz) April 10, 2026 [### Welcome to lab, Elizabeth! ](/news/2026/02/welcome-lab-elizabeth) February 17, 2026 [### Welcome to lab, Fitz! ](/news/2026/01/welcome-lab-fitz) January 09, 2026 arrow\_back arrow\_forward [ More arrow\_circle\_right ](/news) ## Recent Publications Download 3 citations download- [BibTeX](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=bibtex) - [EndNote X3 XML](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=endnote8) - [EndNote 7 XML](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=endnote7) - [Endnote tagged](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=tagged) - [Marc](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=marc) - [PubMedId](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=pubmed_id) - [RIS](/bibcite/export?pager_style=no_pager&number_of_items=3&sort_field=bibcite_year--desc&taxonomy_filters=&&&format=ris) ### 2025 [The developmental basis of floral nectary diversity and evolution](/publication/developmental-basis-floral-nectary-diversity-and-evolution). New Phytologist. 2025;246(6):2462–2477,. doi:https://doi.org/10.1111/nph.70141 [The developmental basis of floral nectary diversity and evolution](/publication/developmental-basis-floral-nectary-diversity-and-evolution). New Phytologist. 2025;246(6):2462–2477,. doi:https://doi.org/10.1111/nph.70141 - add\_circle\_outline do\_not\_disturb\_on Abstract - [ descriptionPublisher's Version](https://doi.org/10.1111/nph.70141) Summary Nectar is a central bridge between angiosperms and animal mutualists. It is produced by specialized structures termed nectaries, which can be found on different plant organs. Consumption of floral nectar by pollinators and the subsequent transfer... - [ descriptionPublisher's Version](https://doi.org/10.1111/nph.70141) ### 2024 Johns J, Min Y, Ballerini E, al. [Loss of staminodes in Aquilegia jonesii reveals a fading stamen-staminode boundary](/publications/loss-staminodes-aquilegia-jonesii-reveals-fading-stamen-staminode-boundary). EvoDevo. 2024;15(6). Johns J, Min Y, Ballerini E, al. [Loss of staminodes in Aquilegia jonesii reveals a fading stamen-staminode boundary](/publications/loss-staminodes-aquilegia-jonesii-reveals-fading-stamen-staminode-boundary). EvoDevo. 2024;15(6). - [ descriptionPublisher's Version](https://rdcu.be/dX0TS) - [ picture\_as\_pdfs13227-024-00225-3.pdf](/sites/g/files/omnuum6406/files/s13227-024-00225-3.pdf) - [ descriptionPublisher's Version](https://rdcu.be/dX0TS) - [ picture\_as\_pdfs13227-024-00225-3.pdf](/sites/g/files/omnuum6406/files/s13227-024-00225-3.pdf) Appleton A, Kramer E. [Genetic Architecture of Novel Floral Organs](/publications/genetic-architecture-novel-floral-organs). International Journal of Plant Science. 2024;185:211–217. Appleton A, Kramer E. [Genetic Architecture of Novel Floral Organs](/publications/genetic-architecture-novel-floral-organs). International Journal of Plant Science. 2024;185:211–217. - [ descriptionPublisher's Version](https://www.journals.uchicago.edu/doi/epdf/10.1086/729359) - [ picture\_as\_pdf729359.pdf](/sites/g/files/omnuum6406/files/729359.pdf) - [ descriptionPublisher's Version](https://www.journals.uchicago.edu/doi/epdf/10.1086/729359) - [ picture\_as\_pdf729359.pdf](/sites/g/files/omnuum6406/files/729359.pdf) [ More arrow\_circle\_right ](/publications)